Seven years after the Battle of Teppelin, humans have reclaimed the earth’s surface and are enjoying an unprecedented era of peace and prosperity. However, the rapid increase in humanity’s population triggers the sudden appearance of an unknown, powerful enemy. The fearsome, manipulative power of the mysterious Anti‑Spiral proves overwhelming, leaving humanity unable to fight back. As hope fades, the members of Team Dai‑Gurren reunite to stand against the threat once more.