During a summer festival five years ago, two girls met beneath a mysterious tree and encountered two glowing spheres. Now, those same girls, Saki Hyūga, the ace pitcher of her school’s softball team, and Mai Mishō, who prefers sketching to stargazing, have been chosen by the spirits of flowers (Flappy) and birds (Choppy) to restore the Seven Fountains and rescue their worlds from Dark Autumn. Together, they become the new Pretty Cure.