Two years after the conclusion of *Martian Successor Nadesico*, the former crew members Akito and Yurika are missing. The Jovians and Earthlings have united, and Ruri has taken command of the newly commissioned Nadesico B. As boson‑jumping technology becomes increasingly popular, a vast transportation network, known as the Hisago Plan, has been established. This system of Chulip portals holds the key to the mysteries of Bose particles and their power. If those secrets were to fall into the wrong hands, the consequences could be dire. A splinter group of Martians launches an attack, leaving the United Forces disoriented and desperate. In response, the former Nadesico crew is recalled to active duty for a special mission.